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Koenig, Walter D
Cornell Academic Staff
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Positions
- Senior Scientist, Cornell Laboratory of Ornithology (CLO)
- Senior Scientist, Neurobiology and Behavior (BIO NB), College of Agriculture and Life Sciences (CALS)
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Affiliations
Research
research overview
- Reproductive strategies of forest trees; spatial synchrony; spatial patterns; conservation biology; behavioral ecology; ornithology; population ecology; vertebrate social behavior; dispersal, mating systems, and sexual selection
principal investigator on
- COLLABORATIVE RESEARCH: REPRODUCTIVE SHARING AND FITNESS IN COOPERATIVE SOCIETIES awarded by DIRECTORATE FOR BIOLOGICAL SCIENCES NSF 2009 - 2013
- LTREB: PHENOLOGY AND MASTING IN CALIFORNIA OAKS awarded by NATL SCIENCE FOUNDATION 2008 - 2013
area(s) of concentration/expertise
keywords
- Behavioral ecology and population biology
Publications
individual publications
-
academic article
- An experimental study of chick provisioning in the cooperatively breeding acorn woodpecker. Ethology. 118:566-574. 2012
- Brooding, provisioning, and compensatory care in the cooperatively breeding acorn woodpecker. Behavioral Ecology. 23:181-190. 2012
- Overlapping landscapes: a persistent, but misdirected concern when collecting and analyzing ecological data. Journal of Wildlife Management. 76:1072-1080. 2012
- Sex allocation in California oaks: trade-offs or resource tracking? . PLoS One. 7. 2012
- Stabilizing selection for within-season flowering phenology confirms pollen limitation in a wind-pollinated tree. Journal of Ecology. 100:758-763. 2012
- Age-related provisioning behaviour in the cooperatively breeding acorn woodpecker: testing the skills and the pay-to-stay hypotheses. Animal Behaviour. 82:437-444. 2011
- Avian predators are less abundant during periodical cicada emergences, but why?. Ecology. 92:784-790. 2011
- Climatic constraints on wintering bird distributions are modified by urbanization and weather. Journal of Animal Ecology. 80:403-413. 2011
- Effects of gypsy moth outbreaks on North American woodpeckers. Condor. 113:352-361. 2011
- Fitness consequences of seed size in the valley oak Quercus lobata Née (Fagaceae). Annals of Forest Science. 68:477-484. 2011
- Fitness consequences of within-brood dominance in the cooperatively breeding acorn woodpecker. Behavioral Ecology and Sociobiology. 65:2229-2238. 2011
- Foraging patterns of acorn woodpeckers (Melanerpes formicivorus) on valley oak (Quercus lobata Née) in two California oak savanna-woodlands. Oceologia. 166:187-196. 2011
- Inter- and intraspecific pollination patterns of valley oak, Quercus lobata, in a mixed stand in central coastal California. International Journal of Plant Science. 172:691-699. 2011
- Mutualism or parasitism? Using a phylogenetic approach to characterize the oxpecker-ungulate relationship. Evolution. 65:1297-1304. 2011
- Proactive Conservation Management of an Island-endemic Bird Species in the Face of Global Change . BioScience. 61:1013-1021. 2011
- Testing Moran’s theorem in an agroecosystem. Oikos. 120:1434-1440. 2011
- Variable helper effects, ecological conditions, and the evolution of cooperative breeding in the acorn woodpecker. American Naturalist. 178:145-158. 2011
- Ecological determinants of American crow mortality due to West Nile virus during its North American sweep. Oecologia. 163:903-909. 2010
- Testing the environmental prediction hypothesis for mast-seeding in California oaks. Canadian Journal of Forest Research. 40:2115-2122. 2010
- Helpers and egg investment in the cooperatively breeding acorn woodpecker: testing the concealed helper effects hypothesis. Behavioral Ecology and Sociobiology. 63:1659-1665. 2009
- Latitudinal decrease in acorn size in bur oak (Quercus macrocarpa) is due to environmental constraints, not avian dispersal. Botany. 87:349-356. 2009
- Mast-producing trees and the geographical ecology of western scrub-jays. Ecography. 32:561-570. 2009
- No trade-off between seed size and number in the valley oak Quercus lobata. American Naturalist. 173:682-688. 2009
- A tale of two worlds: molecular ecology and population structure of the threatened Florida scrub-jay. Molecular Ecology. 17:1632-1633. 2008
- Acorn harvesting by acorn woodpeckers: annual variation and comparison with genetic estimates. Evolutionary Ecology Research. 10:811-822. 2008
- Acorns, insects, and the diet of adult vs. nestling acorn woodpeckers. Journal of Field Ornithology. 79:280-285. 2008
- An intercontinental comparison of the dynamic behavior of mast seeding communities. Population Ecology. 50:329-342. 2008
- Early impact of West Nile virus on the yellow-billed magpie (Pica nuttalli). Auk. 125:542-550. 2008
- Estimate of Trichomonas gallinae-induced mortality in band-tailed pigeons, upper Carmel Valley, California, winter 2006-2007. Wilson Journal of Ornithology. 120:603-606. 2008
- Herbarium specimens suggest non-native origins of three species of Phytophthora in California. Mycological Research. 112:757-758. 2008
- A negative correlation does not imply a tradeoff between growth and reproduction in California oaks. Proceedings at the National Academy of Science USA. 104:16982-16985. 2007
- Costs and benefits of nest reuse versus excavation in cavity-nesting birds. Annales Zoologici Fennici. 44:209-217. 2007
- In Memoriam: Howard L. Cogswell, 1915-2006. Auk. 124:1096. 2007
- Late summer and fall nesting in the acorn woodpecker and other North American terrestrial birds. Condor. 109:334-350. 2007
- Long-term growth and persistence of blue oak (Quercus douglasii) seedlings in a California oak savanna. Madroño. 54:269-274. 2007
- West Nile Virus and breeding bird declines. Ecohealth. 4:18-24. 2007
- Abbreviated inner primaries: a sex-linked dimorphism in the acorn woodpecker. Journal of Field Ornithology. 77:157-162. 2006
- Estimating the effects of sudden oak death on oak-dependent birds. Biological Conservation. 127:146-157. 2006
- Evolution of clutch size in cavity-excavating birds: the nest site limitation hypothesis revisited. American Naturalist. 167:343-353. 2006
- Spatial synchrony of monarch butterflies. American Midland Naturalist. 155:39-49. 2006
- Effects of periodical cicada emergences on abundance and synchrony of avian populations. Ecology. 86:1873-1882. 2005
- Persistence in adversity: lessons from the ivory-billed woodpecker. BioScience. 55:646-647. 2005
- Seasonal body weight variation in five species of woodpeckers. Condor. 107:810-812. 2005
- In Memoriam: Frank Alois Pitelka, 1916-2003. Auk. 121:963-965. 2004
- Spatial synchrony in population dynamics. Annual Review of Ecology, Evolution and Systematics. 35:467-490. 2004
- Desperately seeking similarity. Science. 300:1887-1889. 2003
- Dissecting components of population-level variation in seed production and the evolution of masting behavior. Oikos: A Journal of Ecology. 102:581-591. 2003
- European starlings and their effect on native cavity-nesting birds. Conservation Biology. 17:1134-1140. 2003
- Is pollen limited? The answer is blowin’ in the wind. Trends in Ecology and Evolution. 18:157-159. 2003
- Measuring mast seeding behavior: relationships among population variation, individual variation and synchrony. Journal of Theoretical Biology. 224:107-114. 2003
- Patterns of reproductive skew in the polygynandrous acorn woodpecker. American Naturalist. 162:277-289. 2003
- Regional dynamics of wetland-breeding frogs and toads: turnover and synchrony. Ecological Applications. 13:1522-1532. 2003
- Regional impacts of periodical cicadas on oak radial increment. Canadian Journal of Forest Research. 33:1084-1089. 2003
- Global patterns of environmental synchrony and the Moran effect. Ecography. 25:283-288. 2002
- Limited hybridization between Quercus lobata and Quercus douglasii (Fagaceae) in a mixed stand in central coastal California. American Journal of Botany. 89:1792-1798. 2002
- Reproductive skew in the polygynandrous acorn woodpecker. Proceedings at the National Academy of Science USA. 99:7178-7183. 2002
- Size, insect parasitism, and energetic value of acorns stored by acorn woodpeckers. Condor. 104:539-547. 2002
- Social and genetic monogamy in territorial and loosely colonial populations of the phainopepla. Auk. 119:770-777. 2002
- Dividing up the kids. Science. 291:442-443. 2001
- Nestling sex-ratio variation in the cooperatively breeding acorn woodpecker (Melanerpes formicivorus). Behavioral Ecology and Sociobiology. 49:357-365. 2001
- Seed-crop size and eruptions of North American boreal seed-eating birds. Journal of Animal Ecology. 70:609-620. 2001
- Shared parentage and incest avoidance in the cooperatively breeding acorn woodpecker. Molecular Ecology. 10:1515-1526. 2001
- Spatial autocorrelation and local disappearances in wintering North American birds. Ecology. 82:2636-2644. 2001
- Spatially autocorrelated demography and interpond dispersal in the pond breeding salamander Ambystoma californiense. Ecology. 82:3519-3530. 2001
- Synchrony and periodicity of eruptions by boreal seed-eating birds. Condor. 103:725-735. 2001
- Annual variation in xylem water potential in California oaks. Madroño. 47:106-108. 2000
- Life history and demographic variation in the California tiger salamander (Ambystoma californiense). Copeia Journal. 2000:365-377. 2000
- Natal dispersal in the cooperatively breeding acorn woodpecker. Condor. 102:492-502. 2000
- Patterns of annual seed production by northern hemisphere trees: a global perspective. American Naturalist. 155:59-69. 2000
- Demographic consequences of incest avoidance in the cooperatively breeding acorn woodpecker. Animal Behaviour. 57:1287-1293. 1999
- Nest-site selection in the acorn woodpecker. Auk. 116:45-54. 1999
- Oaks, acorns, and the geographical ecology of the acorn woodpecker. Journal of Biogeography. 26:159-165. 1999
- Sex-ratio selection in species with helpers at the nest: the repayment model revisited. American Naturalist. 153:124-130. 1999
- Spatial autocorrelation in ecological studies. Trends in Ecology and Evolution. 14:22-26. 1999
- Spatial dynamics in the absence of dispersal: acorn production by oaks in central coastal California. Ecography. 22:499-506. 1999
- Synchrony and asynchrony of acorn production at two coastal California sites. Madroño. 46:20-24. 1999
- The ecology of blue oak (Quercus douglasii) acorn production. Journal of the International Oak Society. 9:60-68. 1999
- Effects of storage on tannin and protein content of cached acorns. Southwestern Naturalist. 43:170-175. 1998
- Reproductive roles in the cooperatively breeding acorn woodpecker: incest avoidance vs. reproductive competition. American Naturalist. 151:243-255. 1998
- Scale of mast-seeding and tree-ring growth. Nature. 396:225-226. 1998
- Spatial autocorrelation in California land birds. Conservation Biology. 12:612-620. 1998
- Testing for spatial autocorrelation in ecological studies. Ecography. 21:423-429. 1998
- Host preferences and behavior of oxpeckers: coexistence of similar species in a fragmented landscape. Evolutionary Ecology. 11:91-104. 1997
- Individual, brood, and sex variation in begging calls of western bluebirds. Wilson Bulletin. 109:328-332. 1997
- On the relationship between nutrient use efficiency and fertility in forest ecosystems. Oecologia. 110:550-556. 1997
- Reproductive success of yellow-billed magpies Pica nuttalli in central coastal California. Acta Ornithologica. 32:99-109. 1997
- Site fertility and leaf nutrients of sympatric evergreen and deciduous species of Quercus in central coastal California. Plant Ecology. 130:121-131. 1997
- Acorn production by oaks in central coastal California: influence of weather at three levels. Canadian Journal of Forest Research. 26:1677-1683. 1996
- Detectability, philopatry, and the distribution of dispersal distances in vertebrates. Trends in Ecology and Evolution. 11:514-517. 1996
- Nestling sex-ratio variation in western bluebirds. The Auk. 113:902-910. 1996
- The fitness consequences of helping behavior in the western bluebird. Behavioral Ecology. 7:168-177. 1996
- An experimental test of offspring recognition in western bluebirds. The Auk. 112:1062-1064. 1995
- Genetic monogamy in single-male groups of acorn woodpeckers. Molecular Ecology. 4:765-770. 1995
- Patterns and consequences of egg destruction among joint-nesting acorn woodpeckers. Animal Behaviour. 50:607-621. 1995
- Why do oaks produce boom-and-bust seed crops?. California Agriculture. 49:7-12. 1995
- Acorn production by oaks in central coastal California: variation within and among years. Ecology. 75:99-109. 1994
- Energetic benefits of communal roosting by acorn woodpeckers in the non-breeding season. Condor. 96:631-637. 1994
- Estimating acorn crops using visual surveys. Canadian Journal of Forest Research. 24:2105-2112. 1994
- Provisioning in western bluebirds is not related to offspring sex. Behavioral Ecology. 5:455-459. 1994
- Techniques for capturing birds inside natural cavities. Journal of Field Ornithology. 65:70-75. 1994
- Two new bird-mammal associations from Kenya, with comments on host use by wattled starlings. Ostrich. 65:337-338. 1994
- Water use strategies of five sympatric oaks in central coastal California. Madroño. 41:290-301. 1994
- Boxes, barns, and bridges: confounding factors or exceptional opportunities in ecological studies?. Oikos: A Journal of Ecology. 63:305-308. 1992
- Copulation and mate guarding in the colonial yellow-billed magpie Pica nuttalli and a comparison with the solitary black-billed magpie P. pica. Behaviour. 121:110-130. 1992
- Gopher snake attraction to birds' nests. Southwestern Naturalist. 37:293-298. 1992
- The evolution of delayed dispersal in cooperative breeders. Quarterly Review of Biology. 67:111-150. 1992
- Distress calls in the acorn woodpecker. Condor. 93:637-643. 1991
- Explanations for avian helping behavior. Trends in Ecology and Evolution. 6:343-344. 1991
- Levels of female choice in the white-tailed skimmer Plathemis lydia (Odonata: Libellulidae). Behaviour. 119:193-224. 1991
- The effects of tannins and lipids on the digestibility of acorns by acorn woodpeckers. Auk. 108:79-88. 1991
- Breeding energetics and thermal ecology of the acorn woodpecker in central coastal California. Condor. 92:341-359. 1990
- Dispersal, effective population size and the genetic structure of the contemporary United States. American Journal of Human Biology. 2:165-170. 1990
- Individual contributions to cooperative nest care in the acorn woodpecker. Condor. 92:360-368. 1990
- Opportunity of parentage and nest destruction in polygynandrous acorn woodpeckers, Melanerpes formicivorus. Behavioral Ecology. 1:55-61. 1990
- Territory size and duration in the white-tailed skimmer Plathemis lydia (Odonata: Libellulidae). Journal of Animal Ecology. 59:317-334. 1990
- Helping behaviour, reproductive value, and the future component of indirect fitness. Animal Behaviour. 38:331-343. 1989
- Sex-biased dispersal in the contemporary United States. Ethology and Sociobiology. 10:263-278. 1989
- Ability of two species of oak woodland birds to subsist on acorns. Condor. 90:705-708. 1988
- Costs and benefits of joint nesting in the acorn woodpecker. American Naturalist. 131:654-677. 1988
- Internal migration in the contemporary United States: comparison of measures and partitioning of stages. Human Biology. 60:927-944. 1988
- La vie communicataire chez les oiseaux. La Recherche (Paris). 19:300-308. 1988
- On the determination of viable population size in birds and mammals. Wildlife Society Bulletin. 16:230-234. 1988
- Reciprocal altruism in birds: a critical review. Ethology and Sociobiology. 9:73-84. 1988
- Breeding site fidelity in Plathemis lydia (Anisoptera: Libellulidae). Odonatologica. 16:249-259. 1987
- Lifetime reproductive success, selection, and the opportunity for selection in the white-tailed skimmer Plathemis lydia (Odonata: Libellulidae). Evolution. 41:22-36. 1987
- Morphological and dietary correlates of clutch size in North American woodpeckers. Auk. 104:757-765. 1987
- Poor acorn crop, dominance, and decline in numbers of acorn woodpeckers. Journal of Animal Ecology. 56:197-207. 1987
- Potential poisoning of yellow-billed magpies by compound 1080. Wildlife Society Bulletin. 15:274-276. 1987
- Comparative reproductive success of yellow-shafted, red-shafted and hybrid flickers across a hybrid zone. Auk. 103:42-51. 1986
- Geographic ecology of clutch size variation in North American woodpeckers. Condor. 88:499-504. 1986
- On the measurement of sexual selection. American Naturalist. 127:403-409. 1986
- An analysis of genetic variation and parentage in a California population of acorn woodpeckers. Auk. 102:305-312. 1985
- Dunking of prey by Brewer's blackbirds: a novel source of water for nestlings. Condor. 87:444-445. 1985
- Patterns of territoriality and mating success in the white-tailed skimmer Plathemis lydia (Odonata: Anisoptera). American Midland Naturalist. 114:1-12. 1985
- Replacement of breeders and within-group conflict in the cooperatively breeding acorn woodpecker. Behavioral Ecology and Sociobiology. 17:303-312. 1985
- Clutch size of the gilded flicker. Condor. 86:89-90. 1984
- Geographic variation in clutch size in the northern flicker (Colaptes auratus): support for Ashmole's hypothesis. Auk. 101:698-706. 1984
- The breeding system of the acorn woodpecker in central coastal California. Zeitshrift für Tierpsychologie. 65:289-308. 1984
- Are acorn woodpecker territories aggregated?. Ecology. 64:1305-1307. 1983
- Mate-guarding in the acorn woodpecker: within-group reproductive competition in a cooperative breeder. Animal Behaviour. 31:1094-1106. 1983
- Reproductive competition in the communal acorn woodpecker: sisters destroy each other's eggs. Nature. 306:583-584. 1983
- Ecological and social factors affecting hatchability of eggs. Auk. 99:526-536. 1982
- Intragroup dynamics of a cooperative breeder: an analysis of reproductive roles in the acorn woodpecker. Behavioral Ecology and Sociobiology. 11:195-201. 1982
- Coalitions of male lions: making the best of a bad job?. Nature. 293:413. 1981
- Reproductive success, group size, and the evolution of cooperative breeding in the acorn woodpecker. American Naturalist. 117:421-443. 1981
- Space competition in the acorn woodpecker: power struggles in a cooperative breeder. Animal Behaviour. 29:396-409. 1981
- Birds of Hastings Reservation, Monterey County California. Western Birds. 11:113-128. 1980
- The determination of runt eggs in birds. Wilson Bulletin. 92:103-107. 1980
- The incidence of runt eggs in woodpeckers. Wilson Bulletin. 92:169-176. 1980
- Variation and age determination in a population of acorn woodpeckers. Journal of Field Ornithology. 51:10-16. 1980
- Water dependence of birds in a temperate oak woodland. Auk. 97:339-350. 1980
- Notes on the status of acorn woodpeckers in central Mexico. Condor. 81:317-318. 1979
- Relatedness and inbreeding avoidance: counterploys in the communally nesting acorn woodpecker. Science. 206:1103-1105. 1979
- Characteristics of storage trees used by acorn woodpeckers in two California woodlands. Journal of Forestry. 76:162-164. 1978
-
article
- A look at Sudden Oak Death 2006
- The mystery of masting in trees 2005
- In Memoriam: Frank Alois Pitelka, 1916-2003 2004
- Group sex in the acorn woodpecker: who comes out on top, and why? 2002
- Oaks, acorns, and acorn woodpeckers 2000
- Groepsgedrag van de Eikelspecht 1998
- Geographical ecology of acorn production by California oaks 1997
- The great egg-demolition derby 1997
- Acorn production in California 1995
- Defender of granaries: the acorn woodpecker 1992
- Oaks, acorns, and the acorn woodpecker 1990
- Cooperative breeding in the acorn woodpecker 1984
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book
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booksection
- Social behaviour, animal 2002
- The behavioral ecology of masting in oaks 2002
- Cooperative breeding, evolution of 1998
- Ecological constraint hypothesis 1998
- Natural and sexual selection components of Odonate mating patterns 1997
- Responses of terrestrial vertebrates to habitat edges and corridors 1996
- Acorn woodpecker (Melanerpes formicivorus) 1995
- Acorn woodpecker 1993
- Cannibalism in birds 1992
- The acorn woodpecker 1991
- Acorn woodpeckers: Group living and food storage under contrasting ecological conditions 1990
- Introduction 1990
- Levels of analysis and the functional significance of helping behavior 1990
- Female roles in cooperatively breeding acorn woodpeckers 1983
- Ecological factors and kin selection in the evolution of cooperative breeding in birds 1981
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chapter
- Acorn woodpecker. The Monterey Pine Forest. 17. 2011
- Cooperative breeding as an alternative reproductive tactic. Alternative Reproductive Tactics: an Integrative Approach. 451-470. 2008
- Lifetime reproductive success and sexual selection theory. Dragonflies and Damselflies. 153-167. 2008
- Reproductive skew in avian societies. Reproductive Skew in Vertebrates: Proximate and Ultimate Causes. 227-264. 2008
- Incest avoidance. Ecology and Evolution of Cooperative Breeding in Birds. 142-156. 2004
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conference paper
- Arboreal seed removal and insect damage in three California oaks. Proceedings of the Fifth Symposium on Oak Woodlands: Oaks in California’s Changing Landscape. 2002
- Patterns of geographic synchrony in growth and reproduction of oaks within California and beyond. Proceedings of the Symposium on Oak Woodlands: Ecology, Management, and Urban Interface Issues. 101-108. 1997
- Determinants of acorn productivity among five species of oaks in central coastal California. Proceedings of the Symposium on Oak Woodlands and Hardwood Rangeland Management. 1991
- Parent-offspring conflict in the acorn woodpecker. Proceedings of the International Ornithological Congress. 1221-1230. 1988
- Acorn production by five species of oaks over a seven year period at the Hastings Reservation, Carmel Valley, California. Symposium on Multiple-Use Management of California's Hardwood Resources. 1987
- Acorn storage by acorn woodpeckers in an oak woodland: an energetics analysis. Symposium on the Ecology, Management, and Utilization of California Oaks. 1980
-
report
- Spatial synchrony and temporal patterns in acorn production by California black oaks. 343-356. 2008
- The impact of West Nile virus on birds in California’s hardwood rangelands. 151-163. 2008
- Timing of flowering and seed production in three California oaks. 371-380. 2008
- Potential effects of SOD on the oak woodland bird community of coastal California. 195-209. 2006
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review
- Wytham Woods: Oxford’s Ecological Laboratory, edited by P. S. Savill, C. M. Perrins, K. J. Kirby, and N. Fisher 2010
- Dispersal, edited by J. E. Danchin, A. A. Dhondt, and J. D. Nichols 2002
- The Red-cockaded Woodpecker: Surviving in a Fire-maintained Ecosystem, by R. N. Conner, D. C. Rudolph, and J. R. Walters. 831-832. 2002
- Helpers at Birds’ Nests: a Worldwide Survey of Cooperative Breeding and Related Behavior, by A. F. Skutch. 273-274. 2000
- Hope is the Thing with Feathers, by C. Cokinos. 963-964. 2000
- Fairy-wrens and Grasswrens, by I. Rowley and E. Russell. 88-89. 1999
- Made for Each Other, a Symbiosis of Birds and Pines, by R. N. Lanner. 100. 1998
- Cooperative Breeding in Mammals, edited by N. G. Solomon and J. A. French. 412-413. 1997
- Woodpeckers: a Guide to Woodpeckers of the World, by H. Winkler, D. A. Christie, and D. Nurney. 726-727. 1996
- The Natural History of Inbreeding and Outbreeding, edited by N. W. Thornhill. 797-798. 1994
- The Ostrich Communal Breeding System, by B. C. R. Bertram. 762-763. 1993
- Food Hoarding in Animals, by S. B. Vander Wall. 690. 1991
- Reproductive Success. Studies of Individual Variation in Contrasting Breeding Systems, edited by T. H. Clutton-Brock. 223-224. 1990
- Ecological Aspects of Social Evolution, edited by D. I. Rubenstein and R. W. Wrangham. 1908-1909. 1987
- The Life of the Woodpecker, by A. F. Skutch 1986
- Perspectives in Ornithology, edited by A. H. Brush and G. A. Clark, Jr.. 81-82. 1985
- The Florida Scrub Jay: Demography of a Cooperative-breeding Bird, by G. E. Woolfenden and J. W. Fitzpatrick. 1402-1404. 1985
- Woodpeckers of the World, by L. L. Short. 698-700. 1983
- Introduction 2004
Teaching
teaching overview
- Behavioral ecology and population biology
teaching activities
Service
service to the profession
- International Society for Behavioral Ecology Officer, Treasurer 2000 -
- Cooper Ornithological Society
- Sigma Xi
- National Science Foundation Research Review Panel 2011
- American Ornithologists' Union Council Member 1999 - 2001
- Cooper Ornithological Society Board of Directors 1992 - 1995
- International Society for Behavioral Ecology Officer, Secretary 1992 - 1994
reviewer or editor for
- Botany (Canadian Journal of Botany); Current Zoology; Journal of Animal Ecology; Journal of Ornithology; Journal of Wildlife Management; Oecologia; Oikos; PLoS One; Science; Western North American Naturalist; Wilson Journal of Ornithology
- Ethology
- NERC - Natural Environment Research Council (UK)
- NERC - Natural Environment Research Council (UK)
- National Geographic Society
- National Science Foundation
Background
education and training
- Ph.D. in Zoology, University of California, Berkeley 1978
- B.S. in Biology, Stanford University 1972
awards and honors
- Honorary Member, 2002
- William Brewster Memorial Award, 1999
- Fellow, 1998
- Fellow, 1988
- Elective Member, 1982
Other
college
- CALS
research keyword
- Behavioral ecology and population biology
name prefix
- Dr.